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After fighting on the losing side of a civil war, Captain Malcolm Reynolds (Nathan Fillion) and his crew of the ship Serenity scrape by as a small-time crooks and transportation for hire, laying low on the outskirts of a galaxy ruled by the Universal Alliance. When Reynolds and his crew pick up a young doctor, Simon (Sean Maher), and his telepathic sister, River (Summer Glau) they get a lot more than a couple of passengers. It seems River stumbled onto a secret the Alliance doesn't want anyone to know, and now they'll stop at nothing to bring her in. Pursued on two sides by the Alliance and another harrowing enemy, the Reavers (crazed savages who've descended into cannibalism), Reynolds soon realizes the greatest danger might already be on his ship.